[1]王孝伟a,李澧洪b,赵维俊,等.祁连山青海云杉林冠层结构的空间异质性及其对天然更新的影响[J].山地学报,2025,(5):645-656.[doi:10.16089/j.cnki.1008-2786.000919]
　WANG Xiaoweia,LI Lihongb,ZHAO Weijun,et al.Spatial Heterogeneity of Canopy Structure in Picea crassifolia Forests of the Qilian Mountains and Its Impact on Natural Regeneration[J].Mountain Research,2025,(5):645-656.[doi:10.16089/j.cnki.1008-2786.000919]
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祁连山青海云杉林冠层结构的空间异质性及其对天然更新的影响()
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《山地学报》[ISSN:1008-2186/CN:51-1516]

卷:
期数:
2025年第5期
页码:
645-656
栏目:
山地环境
出版日期:
2025-12-30

文章信息/Info

Title:
Spatial Heterogeneity of Canopy Structure in Picea crassifolia Forests of the Qilian Mountains and Its Impact on Natural Regeneration
文章编号:
1008-2786-(2025)5-645-12
作者:
王孝伟1a; 李澧洪1b; 赵维俊2; 武龙庆2; 张瑞浩3; 王 平1a*
(1.甘肃农业大学 a.资源与环境学院; b.管理学院,兰州 730070; 2. 甘肃省祁连山水源涵养林研究院 祁连山森林生态系统国家定位观测研究站,甘肃 张掖 734000; 3. 河南绿博林业规划设计有限公司,郑州 450003)
Author(s):
WANG Xiaowei1a;  LI Lihong1b;  ZHAO Weijun2;  WU Longqing2;  ZHANG Ruihao3;  WANG Ping1a*
(1. a. College of Resources and Environmental Sciences; b. School of Management, Gansu Agricultural University, Lanzhou 730070, China; 2. State Positioning Observation and Research Station of Forest Ecosystem of the Qilian Mountain, Gansu
关键词:
青海云杉林;  冠层结构;  空间异质性;  天然更新;  祁连山
Keywords:
Picea crassifolia;  canopy structure;  spatial heterogeneity;  natural regeneration;  the Qilian Mountains
分类号:
S718.5
DOI:
10.16089/j.cnki.1008-2786.000919
文献标志码:
A
摘要:
天然更新幼苗、幼树的生长与林冠层结构及其空间异质性密切相关。然而,大样地尺度下山地森林冠层结构空间异质性研究仍显不足,冠层结构对青海云杉林天然更新的影响机制尚不清晰。本研究以祁连山大野口流域动态监测样地青海云杉林(Picea crassifolia)为研究对象,系统测定冠层结构因子(叶面积指数、平均叶倾角、孔隙度)与更新特征(更新苗密度、地径、株高、冠幅),解析冠层结构空间异质性及其对天然更新的影响。研究结果表明,(1)冠层结构因子变异函数最佳理论模型均为指数型; 叶面积指数与平均叶倾角空间结构比>75%,呈强烈空间自相关,孔隙度空间结构比为50.05%,属中等自相关,方向上,叶面积指数与孔隙度呈南—北优势格局,平均叶倾角无显著方向性,空间分布上,叶面积指数与平均叶倾角呈明显斑块分布,孔隙度呈连续带状分布;(2)冠层结构因子存在显著阈值效应,叶面积指数1.34～1.64、平均叶倾角46.64°～61.01°、孔隙度25%～33%为更新苗分布的最适区间,叶面积指数与幼苗密度极显著正相关(P<0.01),但与幼树地径显著负相关(P<0.05),表明高叶面积指数可能通过降低光照抑制幼树径向生长。本研究定量刻画了冠层结构空间异质性及其与天然更新的关联,明确了影响更新的关键冠层参数优势区间,可为青海云杉林生态过程解析与自然更替促进提供科学依据。
Abstract:
The growth of naturally regenerating seedlings and saplings is closely related to forest canopy structure and its spatial heterogeneity. However, few studies had quantified canopy heterogeneity at large-plot scale in mountain forests, and the mechanisms by which canopy structure influences natural regeneration in Qinghai spruce(Picea crassifolia)forests were poorly understood.
This study systematically measured canopy structural parameters(Leaf Area Index [LAI], Mean Leaf Angle [MLA], Canopy Gap Fraction [CGF])and regeneration characteristics(seedling density, ground diameter, height, crown width)within a 10.2 ha dynamic monitoring plot of Picea crassifolia forest in the Dayekou watershed of the Qilian Mountains, China, followd by analysis of the spatial heterogeneity of canopy structure and its impact on natural regeneration.
(1)Key findings revealed spatial heterogeneity patterns: All three canopy parameters were best fitted by exponential variogram models. LAI and MLA exhibited strong spatial autocorrelation(structural ratios >75%), while CGF showed moderate autocorrelation(50.05%). Directional analysis revealed south-north anisotropy for LAI and CGF, but no directional preference for MLA. Spatially, LAI and MLA displayed patchy distributions, whereas CGF formed continuous bands.
(2)Threshold effects: Optimal regeneration occurred within specific ranges: LAI(1.34-1.64), MLA(46.64°-61.01°), and CGF(25%-33%). LAI showed highly significant positive correlation with seedling density(P<0.01)but significant negative correlation with sapling ground diameter(P<0.05), indicating that high LAI may suppress sapling radial growth through reduced light availability.
This study quantifies the spatial heterogeneity of canopy structure and its relationship with natural regeneration, identifying optimal canopy parameter ranges critical for seedling establishment. These findings provide a scientific basis for elucidating ecological processes and promoting natural succession in Picea crassifolia forests.

参考文献/References:

[1] BONGERS F. Methods to assess tropical rain forest canopy structure: An overview [J]. Plant Ecology, 2001, 153: 263-277. DOI: 10.1023/A:1017555605618
[2] YU Min, SUN Jianxin. Effects of forest patch type and site on herb-layer vegetation in a temperate forest ecosystem [J]. Forest Ecology and Management, 2013, 300: 14-20. DOI: 10.1016/j.foreco.2012.12.039
[3] WAN Pan, HE Ruirui. Canopy structure and understory light characteristics of a natural Quercus aliena var. acuteserrata forest in China northwest: Influence of different forest management methods [J]. Ecological Engineering, 2020, 153: 105901. DOI: 10.1016/j.ecoleng.2020.105901
[4] 解丹丹, 苏志尧. 石门台亚热带森林不同演替阶段冠层结构与林下光照特征[J]. 许昌学院学报, 2018, 37(6): 10-14. [XIE Dandan, SU Zhiyao. Characteristics of canopy structure and understory light in subtropical forest community at different successional stages in Shimentai nature reserve [J]. Journal of Xuchang University, 2018, 37(6): 10-14] DOI: 10.3969/j.issn.1671-9824.2018.06.004
[5] 涂晓云, 王克勤, 赵洋毅, 等. 滇中高原不同林龄华山松林冠层对天然降雨雨滴能量特征的影响[J]. 水土保持通报, 2021, 41(3): 40-49.[TU Xiaoyun, WANG Keqin, ZHAO Yangyi, et al. Effects of Pinus Armandii plantation canopy of different age on energy characteristics of natural rainfall in central Yunnan plateau [J]. Bulletin of Soil and Water Conservation, 2021, 41(3): 40-49] DOI: 10.13961/j.cnki.stbctb.2021.03.006
[6] 邓雅丽, 赵新宇, 崔自杰, 等. 中国森林生态系统林冠层降雨截留特征[J]. 生态学报, 2024, 44(7): 2981-2992. [DENG Yali, ZHAO Xinyu, CUI Zijie, et al. Canopy rainfall interception characteristics of forest ecosystems in China [J]. Acta Ecologica Sinica, 2024, 44(7): 2981-2992] DOI: 10.20103/j. stxb.202307041438
[7] 姚伦芳. 赤水河上游柏木林冠层持水特征比较研究[J]. 环保科技, 2020, 26(2): 10-14. [YAO Lunfang. Comparative study on water-holding characteristics of Cupressus funebris forest in the upper reaches of Chishui River [J]. Environmental Protection Science and Technology, 2020, 26(2): 10-14] DOI: 10.3969/j.issn.1674-0254.2020.02.003
[8] 葛瑶, 王振锡. 天山云杉林冠层结构对草本物种多样性的影响[J]. 西北林学院学报, 2020, 35(6): 74-81. [GE Yao, WANG Zhenxi. Effects of canopy structure on herbaceous species diversity in Picea schrenkiana vat. Tianschanica forest [J]. Journal of Northwest Forestry University, 2020, 35(6): 74-81] DOI: 10.3969/j.issn.1001-7461.2020.06.10
[9] 王莉, 林莎, 李远航, 等. 青海大通典型林分冠层结构与林下植被物种多样性关系研究[J]. 西北植物学报, 2019, 39(3): 524-533.[WANG Li, LIN Sha, LI Yuanhang, et al. Relationship between typical canopy structure and understory vegetation diversity in Datong, Qinghai [J]. Northwestern Journal of Botany, 2019, 39(3): 524-533] DOI: 10.7606/j.issn.1000-4025.2019.03.0524
[10] 黄润霞, 贾小容, 刘婷, 等. 亚热带生态公益林冠层结构与林下辐射动态[J]. 西北林学院学报, 2020, 35(1): 28-36. [HUANG Runxia, JIA Xiaorong, LIU Ting, et al. Canopy structure and understory radiation dynamics of subtropical ecological public welfare forest [J]. Journal of Northwest Forestry University, 2020, 35(1): 28-36] DOI: 10.3969/j.issn.1001-7461.2020.01.05
[11] 高晓晓. 秦岭典型林分林冠层对大气降雨水质影响过程研究[D]. 杨凌: 西北农林科技大学, 2021: 1-51. [GAO Xiaoxiao. Study on the influence process of typical forest canopy on rainfall water quality in Qinling Mountains [D]. Yangling: Northwest Agriculture and Forestry University, 2021: 1-51] DOI: 10.27409/d.cnki.gxbnu.2021.001825
[12] 张星元. 广州城市森林群落林冠结构对林下植被的影响[D]. 广州: 华南农业大学, 2018: 1-55.[ZHANG Xingyuan. Influence of canopy structure on understory vegetation of urban forest communities in Guangzhou, China [D]. Guangzhou: South China Agricultural University, 2018: 1-55]
[13] 谭一波, 何琴飞, 郑威, 等. 珠江流域中上游防护林冠层结构对林下植被的影响[J]. 生态学杂志, 2016, 35(12): 3148-3156. [TAN Yibo, HE Qinfei, ZHENG Wei, et al. Effects of canopy structure on understory vegetation in shelterbelt forests along the middle and upper reaches of Pearl River [J]. Chinese Journal of Ecology, 2016, 35(12): 3148-3156] DOI: 10.13292/j.1000-4890.201612.020
[14] 张星元, 张璐, 马丁, 等. 不同冠层结构下的植物生长型与生活型特征[J]. 中南林业科技大学学报, 2018, 38(7): 37-44.[ZHANG Xingyuan, ZHANG Lu, MA Ding, et al. Characteristics of plant growth type and life forms of different canopy structures in subtropical mountainous forest [J]. Journal of Central South University of Forestry Science and Technology, 2018, 38(7): 37-44] DOI: 10.14067/j.cnki.1673-923x.2018.07.007
[15] 鲁君悦, 吴兆飞, 张春雨, 等. 吉林蛟河针阔混交林林层结构对生产力的影响[J]. 生态学报, 2021, 41(5): 2024-2032.[LU Junyue, WU Zhaofei, ZHANG Chunyu, et al. Influence of forest strate structure on productivity of coniferous and broad-leaved mixed forest in Jiaohe, Jilin [J]. Acta Ecologica Sinica, 2021, 41(5): 2024-2032] DOI: 10.5846/stxb202005131220
[16] 方怡然, 潘澜, 薛立. 冰雪灾害后的杉木人工林冠层结构与林下光照及土壤生化特性的关系[J]. 生态环境学报, 2018, 27(4): 609-616. [FANG Yiran, PAN Lan, XUE Li. Relationships between canopy structure and understory light and soil biochemical property in a Cunninghamia lanceolata stand suffering from ice-snow damage [J]. Ecology and Environmental Sciences, 2018, 27(4): 609-616] DOI: 10.16258/j.cnki.1674-5906.2018.04.003
[17] 敬小丽, 杜伟静, 张璐, 等. 南岭山地森林群落冠层结构与立木多度的关系[J]. 中南林业科技大学学报, 2014, 34(5): 59-65.[JING Xiaoli, DU Weijing, ZHANG Lu, et al. Relationship between canopy structure factors and abundance of standing trees in Nanling mountain forest communities [J]. Journal of Central South Forestry University of Science and Technology, 2014, 34(5): 59-65] DOI: 10.14067/j.cnki.1673-923x.2014.05.014
[18] 邱建丽, 李意德, 陈德祥, 等. 森林冠层结构的生态学研究现状与展望[J].广东林业科技, 2008, 24(1): 75-82. [QIU Jianli, LI Yide, CHEN Dexiang, et al. The research progress and the significance of canopy structure in forest ecology [J]. Guangdong Forestry Science and Technology, 2008, 24(1): 75-82] DOI: 10.3969/j.issn.1006-4427.2008.01.016
[19] 勾啸, 孟诗原, 李炜桢, 等. 华南丘陵地区杉木人工林天然更新研究[J]. 林业与环境科学, 2022, 38(5): 70-76. [GOU Xiao, MENG Shiyuan, LI Weizhen, et al. Natural regeneration of Chinese fir plantations in hilly areas of south China [J]. Forestry and Environmental Science, 2022, 38(5): 70-76] DOI: 10.3969/j. issn.1006 -4427. 2022.05.012
[20] PASTUR G J M, PERI P L, CELLINI J M, et al. Canopy structure analysis for estimating forest regeneration dynamics and growth in Nothofagus pumilio forests [J]. Annals of Forest Science, 2011, 68(3): 587-594. DOI: 10.1007/s13595-011-0059-1
[21] 刘子宣, 贾存, 秦志强, 等. 华北落叶松林下光环境对白扦幼树生长的影响[J]. 南京林业大学学报(自然科学版), 2020, 44(6): 111-117.[LIU Zixuan, JIA Cun, QIN Zhiqiang, et al. Effects of light conditions on the growth of understory Picea meyeri sapling in Larix principis-rupprechtii forest [J]. Journal of Nanjing Forestry University(Natural Science Edition),2020, 44(6): 111-117] DOI: 10.3969/j.issn.1000-2006.201909032
[22] 刘炬, 李永宁. 华北落叶松林隙冠层结构和光环境与云杉幼树生长的关系[J]. 东北林业大学学报, 2023, 51(10): 8-12+27. [LIU Ju, LI Yongning. Relationships between canopy structure and light environment over gaps of Larix principis-rupprechtii and growth of Picea asperata [J]. Journal of Northeastern Forestry University, 2023, 51(10): 8-12+27] DOI: 10.13759/j.cnki.dlxb.2023.10.014
[23] 王卓, 范秀华. 阔叶红松林林冠空隙光强异质性对红松幼树生长的影响[J]. 应用生态学报, 2009, 20(5): 1044-1050. [WANG Zhuo, FAN Xiuhua. Effects of light intensity heterogeneity in gaps of broad leaved Korean pine forest in Changbai Mountains on Pinus koraiensis seedlings growth [J]. Chinese Journal of Applied Ecology, 2009, 20(5): 1044-1050] DOI: 10.13287/j.1001-9332.2009.0183
[24] 董伯骞, 黄选瑞, 徐学华, 等. 退化华北落叶松人工林林隙更新特征[J]. 中南林业科技大学学报, 2014, 34(8): 1-8. [DONG Boqian, HUANG Xuanrui, XU Xuehua, et al. Recruitment characteristics of gaps in degraded Larix pricipis-rupprechtii plantation [J]. Journal of Central South University of Forestry and Technology, 2014, 34(8): 1-8] DOI: 10.14067/j.cnki.1673-923x.2014.08.020
[25] 赵维俊, 牛赟, 杜军, 等. 祁连山青海云杉林叶面积指数的空间异质性[J]. 山地学报, 2024, 42(3): 312-319. [ZHAO Weijun, NIU Yun, DU Jun, et al. Spatial heterogeneity of leaf area index of Picea crassifolia forest in the Qilian Mountains, China [J]. Mountain Research, 2024, 42(3): 312-319] DOI: 10.16089/j.cnki.1008-2786.000825
[26] 刘志理, 毕连柱, 宋国华, 等. 典型阔叶红松林叶面积指数的空间异质性[J]. 北京林业大学学报, 2018, 40(11): 1-11. [LIU Zhili, BI Lianzhu, SONG Guohua, et al. Spatial heterogeneity of leaf area index in a typical mixed broadleaved-Korean pine forest in Xiaoxing'an Mountains of northeastern China [J]. Journal of Beijing Forestry University, 2018, 40(11): 1-11] DOI: 10.13332/j.1000-1522.20170468
[27] 姚丹丹, 雷相东, 余黎, 等. 云冷杉针阔混交林叶面积指数的空间异质性[J]. 生态学报, 2015, 35(1): 71-79. [YAO Dandan, LEI Xiangdong, YU Li, et al. Spatial heterogeneity of leaf area index of mixed spruce-fir-deciduous stands in northeast China [J]. Acta Ecologica Sinica, 2015, 35(1): 71-79] DOI: 10.13332/j.1000-1522.20170468
[28] 温远光, 卢文科, 周晓果, 等. 特大冰冻干扰后大明山常绿阔叶林冠层结构空间异质性动态[J]. 广西科学, 2017, 24(2): 150-158. [WEN Yuanguang, LU Wenke, ZHOU Xiaoguo, et al. Spatial heterogeneity dynamics of canopy structure in evergreen broadleaved forest after a severe ice storm disturbance in Damingshan, southern China [J]. Guangxi Science, 2017, 24(2): 150-158] DOI: 10.13656/j.cnki.gxkx.20170125.004
[29] 区余端, 苏志尧. 粤北山地常绿阔叶林自然干扰后冠层结构空间异质性动态[J]. 植物科学学报, 2012, 30(3): 223-229. [QU Yuduan, SU Zhiyao. Spatial heterogeneity dynamics of canopy structure in a montane evergreen broadleaved forest following a natural disturbance in north Guangdong [J]. Plant Science Journal, 2012, 30(3): 223-229] DOI: 10.3724/SP.J.1142.2012.30223
[30] 刘贤德, 赵维俊, 张学龙, 等. 祁连山排露沟流域青海云杉林土壤养分和pH变化特征[J]. 干旱区研究, 2013, 30(6): 1013-1020. [LIU Xiande, ZHAO Weijun, ZHANG Xuelong, et al. Variation of soil nutrient content and pH value under Picea crassifolia foreat in the Pailougou drainage basin in the Qilian Mountains [J]. Arid Zone Research, 2013, 30(6): 1013-1020] DOI: 10.13866/j.azr.2013.06.009
[31] 赵维俊, 王顺利, 孟好军, 等. 祁连山排露沟流域青海云杉种群结构与空间分布格局[J]. 中南林业科技大学学报, 2018, 38(11): 75-80. [ZHAO Weijun, WANG Shunli, MENG Haojun, et al. Population structure and spatial distribution pattern of Picea crassifolia in Pailougou basin, Qilian Mountains [J]. Journal of Central South University of Forestry Science and Technology, 2018, 38(11): 75-80] DOI: 10.14067/j.cnki.1673-923x.2018.11.011
[32] 拓锋, 刘贤德, 黄冬柳, 等. 祁连山大野口流域青海云杉种群数量动态[J]. 生态学报, 2021, 41(17): 6871-6882. [TA Feng, LIU Xiande, HUANG Dongliu, et al. Quantitative dynamics of Picea crassifolia population in Dayekou basin of Qilian Mountains [J]. Acta Ecologica Sinica, 2021, 41(17): 6871-6882] DOI: 10.5846 /stxb 202004010770
[33] 拓锋, 刘贤德, 刘润红, 等. 祁连山大野口流域青海云杉种群空间格局及其关联性[J]. 植物生态学报, 2020, 44(11): 1172-1183.[TA Feng, LIU Xiande, LIU Runhong, et al. Spatial distribution patterns and association of Picea crassifolia population in Dayekou basin of Qilian Mountains, northwestern China [J]. Chinese Journal of Plant Ecology, 2020, 44(11): 1172-1183] DOI: 10.17521/cjpe.2020.0177
[34] 李妍, 李登武, 韩东辰. 贺兰山东坡青海云杉种群动态[J]. 浙江农林大学学报, 2014, 31(1): 50-56.[LI Yan, LI Dengwu, HAN Dongchen. Population dynamics of Picea crassifolia on the eastern slope of Helan Mountain [J]. Journal of Zhejiang Agriculture and Forestry University, 2014, 31(1): 50-56] DOI: 10.11833/ j.issn.2095-0756.2014.01.008
[35] 王坤, 赵维俊. 祁连山青海云杉林动态监测大样地幼苗幼树更新特征及其对地形因子的响应[J]. 山地学报, 2024, 42(5): 623-639. [WANG Kun, ZHAO Weijun. Characteristics of seedling/sapling regeneration and their response to topographic factors in a large dynamic monitoring sample plot of Picea crassifolia forest in the Qilian Mountains, China [J]. Mountain Research, 2024, 42(5): 623-639] DOI: 10.16089/j.cnki.1008-2786.000849
[36] 杨逍虎, 赵维俊, 闫春鸣, 等. 祁连山青海云杉林动态监测样地更新苗和成树格局的空间变异[J].中南林业科技大学学报, 2022, 42(7): 85-95. [YANG Xiaohu, ZHAO Weijun, YAN Chunming, et al. Spatial variation of regeneration patternsin a dynamic monitoring plot of Picea crassifolia forest in the Qilian Mountains [J]. Journal of Central South University of Forestry and Technology, 2022, 42(7): 85-95] DOI: 10.14067/j.cnki.1673-923x.2022.07.010
[37] 郑新奇,吕利娜. 地统计学(现代空间统计学)[M]. 北京: 科学出版社, 2018: 26-60.[ZHENG Xinqi, LYU Lina. Geostatistics(Modern spatial statistics)[M]. Beijing: Science Press, 2018: 26-60]
[38] EICHHORN M P. Pattern reveals process: Spatial organisation of a Kamchatkan stone birch forest [J]. Plant Ecology & Diversity, 2010, 3(3): 281-288. DOI: 10.1080/17550874.2010.528804
[39] 王政权, 王庆成, 李哈滨. 红松老龄林主要树种的空间异质性特征与比较的定量研究[J]. 植物生态学报, 2000, 24(6): 718-723. [WANG Zhengquan, WANG Qingcheng, LI Habin, Characteristics and comparison of spatial heterogeneity of the main species of Korean pine in old growth forests [J]. Journal of Plant Ecology, 2000, 24(6): 718-723] DOI: 10.3321/j.issn:1005-264X.2000.06.013
[40] 张志, 张梦弢, 申欢欢, 等. 关帝山云杉次生林冠层结构对天然更新的影响[J]. 生态学杂志, 2023, 42(5): 1043-1048. [ZHANG Zhi, ZHANG Mengtao, SHEN Huanhuan, et al. Effects of canopy structure on natural regeneration of Picea asperata secondary forest in Guandi Mountain [J]. Chinese Journal of Ecology, 2023, 42(5): 1043-1048] DOI: 10.13292/j.1000-4890.202305.019
[41] 李晖, 杨华, 谢榕. 长白山云冷杉林林隙冠层特征及与幼苗幼树的关系[J]. 北京林业大学学报, 2021, 43(7): 54-62. [LI Hui, YANG Hua, XIE Rong. Canopy characteristics in gaps and its relationship with seedlings and saplings in a spruce-fir forest in the Changbai Mountain area of northeastern China [J]. Journal of Beijing Forestry University, 2021, 43(7): 54-62] DOI: 10.12171/j.1000.1522.20200131
[42] 李效雄, 刘贤德, 赵维俊. 祁连山大野口流域青海云杉种群结构和空间分布格局[J]. 干旱区地理, 2012, 35(6): 960-967. [LI Xiaoxiong, LIU Xiande, ZHAO Weijun. Population structure and spatial distribution pattern of Picea crassifolia in Dayekou basin of Qilian Mountains [J]. Arid Land Geography, 2012, 35(6): 960-967] DOI: 10.13826/j.cnki.cn65-1103/x.2012.06.003
[43] 刘济萌, 刘贤德, 马瑞, 等. 祁连山排露沟流域青海云杉天然更新空间关联性研究[J]. 干旱区资源与环境, 2024, 38(1): 161-170. [LIU Jimeng, LIU Xiande, MA Rui, et al. Spatial association of natural regeneration of Picea crassifolia in Pailugou watershed, Qilian Mountains [J]. Journal of Arid Land Resource and Environment, 2024, 38(1): 161-170] DOI: 10.13448/j.cnki.jalre.2024.018
[44] 李冰, 樊金拴, 车小强. 我国天然云冷杉针阔混交林结构特征、更新特点及经营管理[J]. 世界林业研究, 2012, 25(3): 43-49. [LI Bing, FAN Jinshuan, CHE Xiaoqiang. A review on structural features, regeneration features and management of natural spruce-fir mixed stand of coniferous and broadleaved trees in China [J]. World Forestry Research, 2012, 25(3): 43-49] DOI: 10.13348/j.cnki.sjlyyj.2012.03.014
[45] 贺丹妮, 杨华, 温静, 等. 长白山云冷杉针阔混交林不同林隙下幼苗幼树密度及空间分布[J].应用生态学报, 2020, 31(6): 1916-1922. [HE Danni, YANG Hua, WEN Jing, et al. Density and spatial distribution of seedlings in different gap sizea of a spruce-fir mixed stand in Changbai Mountains, China [J]. Chinese Journal of Applied Ecology, 2020, 31(6): 1916-1922] DOI: 10.13287/j.1001-9332.202006.004

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备注/Memo

备注/Memo:
收稿日期(Received date): 2025- 03-27; 改回日期(Accepted date):2025-10-28
基金项目(Foundation item): 张掖市市级科技计划项目(ZY2024RC16、ZY2023RC10); 国家自然科学基金(32060247); 甘肃省自然科学基金(23JRRG0022)。[Zhangye Municipal Science and Technology Programme(ZY2024RC16、ZY2023RC10); National Natural Science Foundation of China(32060247); Natural Science Foundation of Gansu Province(23JRRG0022)]
作者简介(Biography): 王孝伟(2000-),男,甘肃和政县人,硕士研究生,主要研究方向:森林与土壤生态。[WANG Xiaowei(2000-), male, born in Hezheng, Gansu Province, M.Sc. candidate, research on forest and soil ecology] E-mail: 2027725726@qq.com
更新日期/Last Update: 2025-10-20